https://www.domestika.org/en/courses/565-pixel-art-character-animation-for-video-games

Make your characters come alive in Pyxel Edit frame by frame
The aesthetics pixel art is back to stay with artists such as Daniel Benítez - video game developer and co-founder of Fourattic studio - that keeps her alive in her creations. In this course, I will teach you the techniques and processes to follow to animate a videogame character and its environment in pixel art using the Pyxel Edit tool.
If you have already completed Daniel's previous course, Introduction to character design in pixel art , in this you will go a step further by animating your creations frame by frame and learning your own animation concepts such as timing , anticipation or motion blur and processes such as export and optimization of sprite sheets .